The cuticle is usually highly abundant in cystine residues, which makes a highly cross-linked structure. Morphologically, the cuticle of the animal fibers is an external sheath, which has also its own architecture, being formed of four layers: the epicuticle, the a-layer, the exocuticle, and the endocuticle. Popescu, in Polymer Science: A Comprehensive Reference, 2012 10.16.3.2 Animal Fiber
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